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In the study by Kondo, the amplification frequency is 159th (Kondo et al., 2013)
Metallic or altered tastes can leave people feeling oral discomfort and may cause poor nutrition
The deprotonated forms of Lys and Arg with lower pKbs are much stronger bases than the deprotonated form of His, so at physiological pH, they would always be protonated (unless their local environment lowers their pKa and pKb values)